A Russian nuclear submarine sinks while submerged with a US nuclear submarine nearby. The Russian government learns that the fault for the sinking lies with the Russians, but the officials seek to blame the US anyway. The US submarine leaders know what took place, but they feel reluctant to reveal that, because to do so would disclose an important advantage. After two surprising twists Mac takes a ride aboard a US submarine, and Harm takes one aboard a Russian submarine. The two boats wind up near each other, because the US vessel shadows the Russian one. An extremely dangerous situation develops aboard the Russian boat, and Mac and Harm take part in resolving the problem. Meanwhile a woman ensign in the US Navy graduates with honors in nuclear engineering from a college and from the Naval Nuclear-power School; she then applies but is rejected to attend the Naval Submarine School and to serve aboard a submarine; Mac and Bud advise her.

Representative Latham and another ride an MV-22 Osprey aircraft; just above the sea a minor mishap occurs. Latham holds an inquiry, and Harm and Mac take part. Harriet investigates an old case of innocence. The guys start planning a party.

During a guest sermon a Marine general has made remarks about the Islamic faith; this publicly boils into a court-martial; Mac prosecutes, and Harm defends. Bud investigates a complaint of racial bias against Sturgis, and he resolves it.