All three Camden generations dread the annual visit of Eric's parents, "The Colonel" (John Camden) and Grandma Ruth Camden. The tension between father and son is heightened when both Eric and "The Colonel" want to adopt a homeless and orphaned nine-year-old boy, George. Annie discovers the fate of her lost wedding rings when seeing them on a woman at the store.

A visit from Eric's parents, strict Marines colonel John and his equally military wife, strikes controlled terror among the grandkids, and even Jimmy Moon, who initially discarded Matt's accurate imitation of the usual boyfriend-grilling. They come to present George, the charming, manly boy they adopted from an orphanage. But Eric discovers detective Will Grayson, whose death was faked in a witness protection program, is back in Glenoak after the crime king's death in prison. He claims his son, none other than George, back, as remains his legal right. After a fugue attempt from George, a creative compromise is devised. Matt meanwhile discovers aunt Julie is on the verge of an alcohol relapse.

Mary and Lucy selfishly make Matt feel 'guilty' he hasn't moved out yet, although Annie still hopes to keep him home in a garage apartment. They even get Hank to find him a menial morning job in the hospital cafeteria. But Simon feels desert when Matt takes an apartment with John Hamilton. Eric and Annie worry about the Colonel's attitude to Julie's marriage, but the grandparents have gone soft after an around-the-world trip, as George's crazy hairstyle illustrates. But the stress gives already-overworked Eric a mild heart attack.

Eric threatens Robbie with eviction because he thinks he's keeping secrets about Mary. The house is abuzz with secrets and preparations for an elaborate surprise party for Eric's birthday. Johnny drops Mercy on the Camdens' doorstep after Frankie abandons them. Mike trusts Lucy enough to look after his mother for a day, during which the woman surprises her, both by speaking and by what she says.

Robbie introduces Lucy to his friend Hugh, feeling she should date again, but gets jealous enough to separate them with lies. Matt stands up against the free clinic's staff and gets Hank to overrule Dr. Kent's diagnose-and-operate routine. Feeling incompetent as a waiter at Pete's Simon resigns, but he gets promoted because of his business-planning skills. Cued by Billy via George, Wilson and Mary (who is failing in firefighter training) realize it's time to decide whether to commit to becoming a family.

Robbie and Lucy gather the courage to tell everybody about their affair, only to find they all knew and don't really care. Without the thrill of sneaking, their first date fizzles. Ben Kinkirk and Mary both pass the firefighters course, but their meeting on whether to celebrate with classmates messes up Mary's reconciliation with fiancé Wilson., and Billy's questions make Wilson decide against her plan to elope. Matt discovers that Hank and Julie are planning a divorce, and Eric and Annie try to find out why and see if they can help fix it.