When beautiful-but-vapid model wannabe Deb has a fatal car accident, she suddenly finds herself in front of Heaven’s gatekeeper, Fred, who declares her a self-centered "zero." Outraged, she attempts to persuade Fred to return her to her shallow existence but is accidentally relegated to the body of the recently deceased Jane Bingum. A brilliant, thoughtful and plus-size attorney with a loyal assistant, Teri, Jane has always lived in the shadow of her more comely colleagues whereas Deb has always relied on her external beauty. Now, by a twist of fate and a bolt of divine intervention, Deb must come to terms with inhabiting Jane’s plus-size frame in the ultimate showdown between brains and beauty.

Deb discovers what life is like for plus-sized Jane as she takes on a case defending a cocktail waitress who is fired after she gains 50 lbs. Grayson and Kim represent Chad in a divorce, but the only thing he wants back from his wife is his kidney.

Jane’s past catches up with her when a case from the prior year is up for retrial. Jane has difficultly containing her emotions and is thrown in contempt of the court and ends up behind bars. Kim and Grayson sue a shrink who "cured" their client’s husband of his multiple personalities, only she got rid of the wrong one.

Jane defends her - rather Deb’s - mom in a divorce suit when Grayson turns down her mom’s request since he is already representing Deb’s dad. Deb gets a chance to spend time with her mom Bobbi and starts to learn to let go of her old life. Kim represents a dog owner who sues a pet cloning company for breach of contract as he feels this clone is NOT the same dog as his first dog, who was a Westminster champion.

Jane is star struck when she gets to save Deb’s favorite fashion model from a tumble with the law, but soon discovers her idol is in more trouble than she thought. Grayson and Kim take on a pro-bono case where a woman refuses to sell her house to developers because it holds too many memories of her deceased husband. asn’t been in the courtroom in a long time.

Jane has to help a man attempt to regain custody, while still dealing with amnesia. Kim and Parker assist an author by figuring out whether she can retain control of the fictional character or if Jonathan Martin (pseudonym the character was created in) has control of it. Kim finds out that Fred is her assistant and Grayson asks every one of the office to pause for a moment and remember Deb's birthday

Jane brings in an age discrimination case only to have Parker hand it over to Kim. Grayson takes on a real estate case where his clients claim their house is haunted and gets closer to Vanessa. Fred decides to help Stacy with her money woes by hiring her to serve divorce papers.

Jane works a case involving cyber bullying; Grayson meets Vanessa's parents; and Kim takes on a divorce case for Parker's former partner and discovers her client has ulterior motives.

Jane finds herself with a double dose of mother woes when Jane’s mom, Elaine, is arrested for exiting an airplane via the emergency slide after being stuck on a tarmac for six hours...

Owen and Kim try to talk to Jane about her relationship with Ian; Teri realizes that Ian is responsible for ruining her chances at stardom; Owen and Jane find themselves on opposite ends of a case.