Emily takes up the sleuthing reins and hunts for answers about Ezra and Mona; Hanna begins a new search after being confronted by Holbrook and Tanner; Spencer and Aria lean on new men in their lives to recover.

It is a rough time for all of the Liars as Spencer is reeling about the memories from her "lost summer," Aria is still nursing a broken heart, Emily is reeling from Paige's betrayal and Hanna unsuccessfully attempts to wade back into the dating pool. But as each girl tries to deal with her individual issues, they must focus their attention on their suspicions surrounding Mrs. DiLaurentis. Assuming that Mrs. D knows more about Ali and the night she went missing, girls decide to help out at her charity bridal fashion show to gain access to the DiLaurentis house to search for answers. Meanwhile, Ella and Jason return to Rosewood, each with something to hide.

Mona returns to Rosewood in a very fragile state, and Hanna is determined to help her; Emily tries to help Sara; Spencer finds help from someone from her past.

The liars are on the hunt for proof that Lesli is "A", but as they ramp up their investigation, the threats escalate.