The city anxiously anticipates the arrival of Heather Taffet, more notoriously known as "The Gravedigger," for her final appeal after being sentenced to death for a kidnap-murder and a series of similar coldblooded crimes. But when a bullet targeted at The Gravedigger is fired from a distance and hits her, the Jeffersonian team is tasked to piece together the sniper's precise location as well as identify the suspect and his motives. With the help of probing investigator Caroline Julian, Booth and Brennan put together a list of likely suspects, but the evidence and the remarkable skill needed to pull off the crime leads the team to believe that the crime was an inside, professional job, leaving Brennan's father Max and even Booth as plausible suspects. Meanwhile, Sweets attempts to put on a brave face after being caught in the firestorm just moments following a chilly encounter with The Gravedigger.

After notoriously murdering "The Gravedigger," sniper-on-the-loose Jacob Broadsky strikes again. This time, his victim is a prior suspect who escaped the justice system. More determined than ever to catch Broadsky, Booth uses his own military training to beat the sniper at his own game. He vows to go to any lengths necessary to ensure that justice is served. In a heated and dramatic face-off, Booth has minutes to catch Broadsky before he hits an intended target, but Broadsky's skill and speed may prove to be a worthy adversary. Meanwhile, Angela's father visits the Jeffersonian and insists on naming his unborn grandchild.

The search for Broadsky intensifies as he manages to kill one of the Squints. Angela and Hodgins name their baby. Booth struggles to protect those closest to him, which leads to an unexpected situation.